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Protein / target

Lanosterol 14-alpha demethylase

ERG11P10613Candida albicans (strain SC5314 / ATCC MYA-2876)Swiss-Prot
30
Clinical trials
5
Drugs targeting

Protein at a glance

Biological role

Sterol 14-demethylase

Derived from structured UniProt, Open Targets and literature data on this page.

Protein profile

UniProt 2026_02

Canonical identity and biological annotation from UniProt.

Function overview

Sterol 14alpha-demethylase that plays a critical role in the third module of ergosterol biosynthesis pathway, being ergosterol the major sterol component in fungal membranes that participates in a variety of functions.

View complete UniProt function annotation

Sterol 14alpha-demethylase that plays a critical role in the third module of ergosterol biosynthesis pathway, being ergosterol the major sterol component in fungal membranes that participates in a variety of functions (PubMed:10393548, PubMed:28258218, PubMed:8647850, PubMed:9559662). The third module or late pathway involves the ergosterol synthesis itself through consecutive reactions that mainly occur in the endoplasmic reticulum (ER) membrane (By similarity). In filamentous fungi, during the initial step of this module, lanosterol (lanosta-8,24-dien-3beta-ol) can be metabolized to eburicol (By similarity). Sterol 14alpha-demethylase catalyzes the three-step oxidative removal of the 14alpha-methyl group (C-32) of both these sterols in the form of formate, and converts eburicol and lanosterol to 14-demethyleburicol (4,4,24-trimethylergosta-8,14,24(28)-trienol) and 4,4-dimethyl-5alpha-cholesta-8,14,24-trien-3beta-ol, respectively, which are further metabolized by other enzymes in the pathway to ergosterol (PubMed:10393548, PubMed:28258218, PubMed:8647850, PubMed:9559662). Can also use substrates not intrinsic to fungi, such as 24,25-dihydrolanosterol (DHL), producing 4,4-dimethyl-8,14-cholestadien-3-beta-ol, but at lower rates than the endogenous substrates (By similarity)

Subcellular location

Endoplasmic reticulum membrane
Domains and Gene Ontology detail (14)

Gene Ontology

  • Ccortical endoplasmic reticulum
  • Cendoplasmic reticulum membrane
  • Cmembrane
  • Cperinuclear endoplasmic reticulum
  • Cplasma membrane
  • Fheme binding
  • Firon ion binding
  • Foxidoreductase activity
  • Fsterol 14-demethylase activity
  • Pcell growth mode switching, budding to filamentous
  • Pcellular response to xenobiotic stimulus
  • Pendosome organization

528 aa · 61 kDa

Biological roles

What this protein does, drawn together from its UniProt function, Gene Ontology terms and Reactome pathways.

Lipid & lipoprotein metabolismUniProt · GO
View supporting evidence

Lipid & lipoprotein metabolism

  • ·Sterol 14alpha-demethylase that plays a critical role in the third module of ergosterol…
  • ·sterol 14-demethylase activity
  • ·ergosterol biosynthetic process

Concepts derived from UniProt GO Reactome — each badge above shows which sources supported that role.

Approved medicines with mapped indications

5 medicines · 11 areas

Approved therapies targeting this protein, grouped by what they are approved to treat — the disease-first view of the medicines below. Relationships come from the canonical approved-indication graph (ChEMBL phase-4), the same source as the drug cards.

Aspergillosis3 medicines
Infections2 medicines
Candidemia1 medicine
Candidiasis, Vulvovaginal1 medicine
Lung Diseases1 medicine
Neutropenia1 medicine
Pruritus1 medicine
Tinea1 medicine
Urinary Tract Infections1 medicine

Approved indications from ChEMBL (phase-4), via the canonical drug→disease graph.

Drugs targeting this protein

5

How approved and investigational drugs engage this protein — mechanism and action type, direct vs complex targeting, and how broadly each acts across other targets. A drug-first view (the section above groups the approved ones disease-first); direct binders with few recorded targets are listed first.

Voriconazole
Narrow target profileInhibitor

Cytochrome P450 51 inhibitor

Indicated for Aspergillosis, Candidemia, Candidiasis, Infections

Direct interaction with this protein · Only this protein recorded as a target

fluconazole
Narrow target profileInhibitor

Cytochrome P450 51 inhibitor

Indicated for Candidiasis, Candidiasis, Vulvovaginal, Urinary Tract Infections

Direct interaction with this protein · Only this protein recorded as a target

itraconazole
Narrow target profileInhibitor

Cytochrome P450 51 inhibitor

Indicated for Aspergillosis, Candidiasis, Lung Diseases, Mycoses

Direct interaction with this protein · Only this protein recorded as a target

miconazole
Narrow target profileInhibitor

Cytochrome P450 51 inhibitor

Indicated for Infections, Mycoses, Pruritus, Tinea

Direct interaction with this protein · Only this protein recorded as a target

posaconazole
Narrow target profileInhibitor

Cytochrome P450 51 inhibitor

Indicated for Aspergillosis, Candidiasis, Mycoses, Neutropenia

Direct interaction with this protein · Only this protein recorded as a target

ChEMBL mechanism, action type, target identity and approved indications. Open Targets clinical status.

Clinical trials

30

Trials of drugs that target this protein — reached indirectly through those drugs, so a trial listed here studies the drug, not the protein. Ranked by active status, then clinical phase and recency, and spread across the targeting drugs.

View all trials (26)

UNKNOWN · via fluconazole · NCT00537888

ClinicalTrials.gov via the drug-target graph.

What's happening now

12

Recent therapeutic activity around this target — regulatory actions, clinical trials and safety signals for 5 drugs that target this protein, plus publications where such a drug is a genuine subject. Every item is reached indirectly through the drug, not the protein itself; incidental mentions (a paper that merely measures a drug) and press items are excluded.

  1. Label change2026-09-14

    Label change: ITRACONAZOLE (ANDA205080)

    fda · regulatory · fda · via itraconazole

  2. Label change2026-09-14

    Label change: ITRACONAZOLE (ANDA206741)

    fda · regulatory · fda · via itraconazole

  3. Label change2026-09-11

    Label change: ITRACONAZOLE (ANDA208481)

    fda · regulatory · fda · via itraconazole

  4. Trial status changed2026-09-03

    An Open-label, Phase 1 Study to Individually Characterize the Food Effect, the Drug-Drug Interaction Effect of OAT Inhibition, and the Drug-Drug Interaction Effect of P-glycoprotein Inhibition on the Pharmacokinetics of MK-8527 in Healthy Adult Participants

    Status changed to Active, not recruiting · ClinicalTrials.gov · via itraconazole

  5. Trial status changed2026-08-27

    A Phase 1 Crossover Design Study to Assess the Effect of CYP3A Moderate and Strong Inhibitors (Fluconazole and Itraconazole) and Strong Inducer (Carbamazepine) on the Single-dose Pharmacokinetics of ASP3082 in Healthy Adults

    Status changed to Active, not recruiting · ClinicalTrials.gov · via fluconazole

  6. Trial status changed2026-08-25

    A Phase 1, Open-Label, Fixed-Sequence Study to Investigate the Effect of Itraconazole on the Pharmacokinetics of LY4395089 in Healthy Participants

    Status changed to Completed · ClinicalTrials.gov · via itraconazole

  7. Trial status changed2026-08-06

    A Two-Part Clinical Study to Evaluate the Effects of Multiple Doses of Itraconazole on the Single-Dose Pharmacokinetics of MK-2828 (Part 1) and Multiple Doses of MK-2828 on the Single-Dose PK of Midazolam (Part 2) in Healthy Participants

    Status changed to Active, not recruiting · ClinicalTrials.gov · via itraconazole

  8. Regulatory approval2026-06-29

    Approval: POSACONAZOLE (ANDA213331)

    fda · regulatory · fda · via posaconazole

  9. New publication2020-12-14
    Defining and managing COVID-19-associated pulmonary aspergillosis: the 2020 ECMM/ISHAM consensus criteria for research and clinical guidance.

    The Lancet. Infectious diseases · 2021 · 672 citations · Europe PMC · via Voriconazole

  10. Safety communication2017-09-26

    Drug Safety Update: Miconazole (Daktarin): over-the-counter oral gel contraindicated in patients taking warfarin

    mhra · safety · mhra · via miconazole

  11. Safety communication2016-09-15

    Drug Safety Update: Posaconazole (Noxafil): tablets and oral suspension are not directly interchangeable

    mhra · safety · mhra · via posaconazole

  12. Safety communication2014-12-11

    Drug Safety Update: Voriconazole: reminder of risk of liver toxicity, phototoxicity, and squamous cell carcinoma

    mhra · safety · mhra · via Voriconazole

Objective event titles are shown unmodified; the event kind and significance line are derived from structured fields. Forefront AttentionEvent stream aggregating Europe PMC Regulatory filings ClinicalTrials.gov.