Protein / target

Programmed cell death 1 ligand 1

CD274Q9NZQ7Homo sapiensSwiss-Prot
Clinically validated
Therapeutic maturity
5
Approved medicines
30
Clinical trials
Small-molecule tractable
Druggability
Structure with Ligand

Protein at a glance

Biological role

Transcription coactivator activity

Primary system

Immune system

Strongest disease association

neoplasm

Genetic evidence · score 0.19

Therapeutic maturity

Clinically validated target

5 approved medicines against this target

Druggability

Small molecule

Open Targets tractability · Structure with Ligand

Clinical development

5 approved · 8 in clinical development

30 linked trials

Research activity

Extensively researched

158 papers · latest 2025

Derived from structured UniProt, Open Targets and literature data on this page.

Protein profile

UniProt 2026_02

Canonical identity and biological annotation from UniProt.

Function

Plays a critical role in induction and maintenance of immune tolerance to self (PubMed:11015443, PubMed:28813410, PubMed:28813417, PubMed:31399419). As a ligand for the inhibitory receptor PDCD1/PD-1, modulates the activation threshold of T-cells and limits T-cell effector response (PubMed:11015443, PubMed:28813410, PubMed:28813417, PubMed:36727298). Through a yet unknown activating receptor, may costimulate T-cell subsets that predominantly produce interleukin-10 (IL10) (PubMed:10581077). Can also act as a transcription coactivator: in response to hypoxia, translocates into the nucleus via its interaction with phosphorylated STAT3 and promotes transcription of GSDMC, leading to pyroptosis (PubMed:32929201)

Subcellular location

Cell membraneEarly endosome membraneRecycling endosome membraneNucleusEndomembrane systemSecreted
Domains and Gene Ontology detail (35)

Domains & features

Ig-like V-typeIg-like C2-type

Gene Ontology

  • Ccytosol
  • Cearly endosome membrane
  • Cendoplasmic reticulum membrane
  • Cexternal side of plasma membrane
  • Cextracellular exosome
  • CGolgi membrane
  • Cnuclear speck
  • Cnucleoplasm
  • Cplasma membrane
  • Crecycling endosome membrane
  • Freceptor ligand activity
  • Ftranscription coactivator activity

290 aa · 33 kDa · 4 isoforms

Biological roles

Reactome v97

What this protein does, drawn together from its UniProt function, Gene Ontology terms and Reactome pathways.

Immune signallingUniProt · GOTranscriptional regulationUniProt · GOApoptosis & cell deathGO
View supporting evidence

Immune signalling

  • ·Plays a critical role in induction and maintenance of immune tolerance to self (PubMed:1…
  • ·adaptive immune response
  • ·immune response
  • ·negative regulation of activated T cell proliferation

Transcriptional regulation

  • ·Plays a critical role in induction and maintenance of immune tolerance to self (PubMed:1…
  • ·transcription coactivator activity

Apoptosis & cell death

  • ·positive regulation of activated CD8-positive, alpha-beta T cell apoptotic process
View underlying pathways (7)

Concepts derived from UniProt GO Reactome — each badge above shows which sources supported that role.

Interaction neighbourhood

STRING v12.0

Proteins with the strongest functional or physical association. Node size and line weight reflect STRING confidence; hover or select a partner to inspect one association.

CD80CTLA4PDCD1HAVCR2CD28PDCD1L…ICOSSTAT3LAG3TIGITCD274

10 strongest partners — larger node and heavier line mean higher confidence

Functional and physical associations from STRING v12.0. Only associations with this protein are drawn — partner-to-partner links are not part of this evidence.

Drugs targeting this protein

3

Whether each drug engages this protein directly or through a complex, and how many other targets are recorded for it. Direct binders with few recorded targets are listed first.

atezolizumab
Narrow target profileApprovedInhibitor

Programmed cell death 1 ligand 1 inhibitor

Appears in clinical studies involving lung cancer, urothelial carcinoma, non-small cell lung carcinoma, breast cancer

Direct interaction with this protein · Only this protein recorded as a target

durvalumab
Narrow target profileApprovedInhibitor

Programmed cell death 1 ligand 1 inhibitor

Appears in clinical studies involving non-small cell lung carcinoma, small cell lung carcinoma, urothelial carcinoma, non-small cell lung carcinoma

Direct interaction with this protein · Only this protein recorded as a target

avelumab
Narrow target profileApprovedOther

Programmed cell death 1 ligand 1 other

Appears in clinical studies involving Merkel cell skin cancer, Merkel cell skin cancer, urothelial carcinoma, neuroendocrine neoplasm

Direct interaction with this protein · Only this protein recorded as a target

ChEMBL mechanism, action type and target identity. Open Targets clinical status and indications.

Translational evidence

Open Targets 26

Why this target matters therapeutically — disease associations, drugs in development, tractability and safety, from Open Targets.

Strongest genetic associations

Human genetic evidence — the most direct causal link between this target and a disease.

neoplasm0.19

Genetic · overall 0.59

Highest-confidence therapeutic associations

Diseases where a drug acting on this target has already reached clinical development.

non-small cell lung carcinoma0.98

Clinical · overall 0.68

urothelial carcinoma0.94

Clinical · overall 0.59

Merkel cell skin cancer0.93

Clinical · overall 0.64

small cell lung carcinoma0.92

Clinical · overall 0.62

hepatocellular carcinoma0.87

Clinical · overall 0.61

Highest overall evidence

Remaining associations by Open Targets' aggregated evidence score.

head and neck squamous cell carcinoma0.59

Literature

breast cancer0.55

Literature

lung cancer0.52

Literature

esophageal squamous cell carcinoma0.48

Literature

Show all associations
non-small cell lung carcinoma0.68
Merkel cell skin cancer0.64
small cell lung carcinoma0.62
hepatocellular carcinoma0.61
head and neck squamous cell carcinoma0.59
neoplasm0.59
urothelial carcinoma0.59
breast cancer0.55
lung cancer0.52
esophageal squamous cell carcinoma0.48

Open Targets ranks 1,982 associations for this target and we store the top 10 by aggregated score. The long tail is largely literature co-mention and RNA-expression evidence, not evidence that this target drives those diseases.

Known drugs · 13 total

ATEZOLIZUMABApproval

lung cancer · urothelial carcinoma · non-small cell lung carcinoma

ENVAFOLIMABPhase 3

biliary tract cancer · rectum adenocarcinoma · non-small cell lung carcinoma

BINTRAFUSP ALFAPhase 3

non-small cell lung carcinoma · biliary tract cancer · non-small cell lung carcinoma

SUGEMALIMABApproval

non-small cell lung carcinoma · non-small cell lung carcinoma · neoplasm

ADEBRELIMABPhase 3

small cell lung carcinoma · non-small cell lung carcinoma · colorectal cancer

AVELUMABApproval

Merkel cell skin cancer · Merkel cell skin cancer · urothelial carcinoma

DANBURSTOTUGPhase 2

biliary tract cancer · diffuse large B-cell lymphoma · mature T-cell and NK-cell non-Hodgkin lymphoma

PACMILIMABPhase 2

medulloblastoma · neoplasm · breast cancer

DURVALUMABApproval

non-small cell lung carcinoma · small cell lung carcinoma · urothelial carcinoma

SOCAZOLIMABPhase 3

small cell lung carcinoma · cervical cancer · urothelial carcinoma

Tractability

SM · Structure with LigandSM · High-Quality LigandSM · Med-Quality PocketAB · Approved DrugAB · UniProt loc high confAB · GO CC high confAB · UniProt SigP or TMHMMPR · LiteraturePR · UniProt UbiquitinationPR · Small Molecule BinderOC · Advanced Clinical

Clinical trials

30

Trials of drugs that target this protein — reached indirectly through those drugs, so a trial listed here studies the drug, not the protein.

Show remaining trials (24)

ClinicalTrials.gov via the drug-target graph.

Research activity

158 papers · to 2025

Papers linked directly to this protein. This is the protein's own literature — descriptor-derived papers are kept separate below.

Most cited

Borghaei H · The New England journal of medicine · 2015

Keir ME · Annual review of immunology · 2008

Ferris RL · The New England journal of medicine · 2016

Socinski MA · The New England journal of medicine · 2018

Ayers M · The Journal of clinical investigation · 2017

Recent

Europe PMC papers linked directly to this protein.

Forefront confidence

Synthesis

Our own weighting of the evidence behind each disease association. Human genetics and clinical evidence count for most; literature co-mention counts for little, because two entities sharing an abstract is not evidence that one drives the other.

non-small cell lung carcinomaWell supported
0.84
agreement 0.730.96
Clinical61%Somatic mutation25%Literature13%RNA expression2%

Open Targets aggregate 0.68 · 4 independent evidence families

Merkel cell skin cancerWell supported
0.81
agreement 0.690.93
Clinical62%Somatic mutation26%Literature12%

Open Targets aggregate 0.64 · 3 independent evidence families

hepatocellular carcinomaWell supported
0.79
agreement 0.670.91
Clinical60%Somatic mutation27%Literature14%

Open Targets aggregate 0.61 · 3 independent evidence families

small cell lung carcinomaWell supported
0.79
agreement 0.670.91
Clinical67%Somatic mutation19%Literature14%

Open Targets aggregate 0.62 · 3 independent evidence families

head and neck squamous cell carcinomaWell supported
0.78
agreement 0.660.90
Clinical59%Somatic mutation28%Literature14%

Open Targets aggregate 0.59 · 3 independent evidence families

The evidence agreement range shows how closely the independent evidence families agree — it is not a statistical confidence interval, and nothing here is fitted to outcome data. Derived from Open Targets evidence types under Forefront weighting; the underlying per-type scores are shown above so the calculation can be checked.

What's happening now

57

Recent activity around this target, drawn from one canonical event stream. Every item is reached through 3 drugs that target this protein, so each event is news about that drug rather than about the protein directly.

  1. Industry development2026-04-02
    Another AstraZeneca Emerald glimmers as Imfinzi, Imjudo delay liver cancer progression

    Fierce Pharma · news · via durvalumab

  2. New publication2025-09-08
    Modulation of fibronectin extracellular matrix enhances anti-tumor efficacy of immune checkpoint blockade.

    Cell reports. Medicine · 2025 · Europe PMC · via atezolizumab

  3. New publication2025-08-01
    Early ctDNA and Survival in Metastatic Colorectal Cancer Treated With Immune Checkpoint Inhibitors: A Secondary Analysis of the SAMCO-PRODIGE 54 Randomized Clinical Trial.

    JAMA oncology · 2025 · 9 citations · Europe PMC · via avelumab

  4. New publication2025-01-06
    Autogene cevumeran with or without atezolizumab in advanced solid tumors: a phase 1 trial.

    Nature medicine · 2025 · 66 citations · Europe PMC · via atezolizumab

  5. New publication2024-08-01
    Plasma versus Tissue Tumor Mutational Burden as Biomarkers of Durvalumab plus Tremelimumab Response in Patients with Metastatic Colorectal Cancer in the CO.26 Trial.

    Clinical cancer research : an official journal of the American Association for Cancer Research · 2024 · 15 citations · Europe PMC · via durvalumab

  6. New publication2024-03-29
    CONTACT-01: A Randomized Phase III Trial of Atezolizumab + Cabozantinib Versus Docetaxel for Metastatic Non-Small Cell Lung Cancer After a Checkpoint Inhibitor and Chemotherapy.

    Journal of clinical oncology : official journal of the American Society of Clinical Oncology · 2024 · 48 citations · Europe PMC · via atezolizumab

  7. New publication2024-02-13
    Biomarker-directed targeted therapy plus durvalumab in advanced non-small-cell lung cancer: a phase 2 umbrella trial.

    Nature medicine · 2024 · 88 citations · Europe PMC · via durvalumab

  8. New publication2024-02-02
    Phase 1/2 study of monalizumab plus durvalumab in patients with advanced solid tumors.

    Journal for immunotherapy of cancer · 2024 · 42 citations · Europe PMC · via durvalumab

  9. New publication2024-02-01
    Atezolizumab in Combination With Carboplatin and Survival Outcomes in Patients With Metastatic Triple-Negative Breast Cancer: The TBCRC 043 Phase 2 Randomized Clinical Trial.

    JAMA oncology · 2024 · 30 citations · Europe PMC · via atezolizumab

  10. New publication2024-02-01
    First-Line Ipatasertib, Atezolizumab, and Taxane Triplet for Metastatic Triple-Negative Breast Cancer: Clinical and Biomarker Results.

    Clinical cancer research : an official journal of the American Association for Cancer Research · 2024 · 35 citations · Europe PMC · via atezolizumab

  11. New publication2023-11-01
    Anti-TIGIT Antibody Tiragolumab Alone or With Atezolizumab in Patients With Advanced Solid Tumors: A Phase 1a/1b Nonrandomized Controlled Trial.

    JAMA oncology · 2023 · 53 citations · Europe PMC · via atezolizumab

  12. New publication2023-11-01
    Neoadjuvant Durvalumab Alone or Combined with Novel Immuno-Oncology Agents in Resectable Lung Cancer: The Phase II NeoCOAST Platform Trial.

    Cancer discovery · 2023 · 88 citations · Europe PMC · via durvalumab

Objective event titles are shown unmodified; the event kind and significance line are derived from structured fields. Forefront AttentionEvent stream aggregating Europe PMC Regulatory filings ClinicalTrials.gov.